diff --git a/components/email/email-composer.tsx b/components/email/email-composer.tsx
index a65bc5986..8353eb3d3 100644
--- a/components/email/email-composer.tsx
+++ b/components/email/email-composer.tsx
@@ -538,17 +538,25 @@ export function EmailComposer({
     identities.map(i => i.email),
     [composerAccount?.email, composerAccount?.username, replyAccount?.email, replyAccount?.username],
   );
-  const ownIdentityEmailsKey = ownIdentityEmails.map((e) => e.toLowerCase()).join(',');
+  // "Self-sent" means sent by the account holding the original. A message
+  // another connected account sent to this one was received here, and its
+  // reply must go back to that sender, not to this account's own address.
+  const sourceAccount = replyAccount ?? composerAccount;
+  const sourceAccountEmails = ownReplyAddresses(
+    sourceAccountIdentities.map(i => i.email),
+    [sourceAccount?.email, sourceAccount?.username],
+  );
+  const ownIdentityEmailsKey = [...ownIdentityEmails, '|', ...sourceAccountEmails].map((e) => e.toLowerCase()).join(',');
 
   // Initialize with reply/forward data if provided
   const getInitialTo = (): Recipient[] => {
     if (mode !== 'reply' && mode !== 'replyAll') return [];
-    return buildReplyRecipients(replyTo, mode, ownIdentityEmails).to.map(toRecipient);
+    return buildReplyRecipients(replyTo, mode, ownIdentityEmails, sourceAccountEmails).to.map(toRecipient);
   };
 
   const getInitialCc = (): Recipient[] => {
     if (mode !== 'replyAll') return [];
-    return buildReplyRecipients(replyTo, mode, ownIdentityEmails).cc.map(toRecipient);
+    return buildReplyRecipients(replyTo, mode, ownIdentityEmails, sourceAccountEmails).cc.map(toRecipient);
   };
 
   const getInitialSubject = () => {
@@ -1194,10 +1202,10 @@ export function EmailComposer({
     // identity that sent it. Resolving from the recipients here would pick the
     // *other* party's address - and on a catch-all domain it would even set a
     // From override to their address. An identity on the message's own account
-    // wins when two accounts share the address.
-    if (isSelfSent({ from: replyTo?.from }, identities.map(i => i.email).filter(Boolean))) {
-      const senderIdentityId = findDraftIdentityId(sourceAccountIdentities, replyTo?.from?.[0])
-        ?? findDraftIdentityId(identities, replyTo?.from?.[0]);
+    // wins when two accounts share the address. Only the message's own account
+    // counts: mail another connected account sent here is a received message.
+    if (isSelfSent({ from: replyTo?.from }, sourceAccountIdentities.map(i => i.email).filter(Boolean))) {
+      const senderIdentityId = findDraftIdentityId(sourceAccountIdentities, replyTo?.from?.[0]);
       if (senderIdentityId) {
         setSelectedIdentityId(senderIdentityId);
         return;
diff --git a/lib/__tests__/reply-recipients.test.ts b/lib/__tests__/reply-recipients.test.ts
index c3966e514..14ef07ec8 100644
--- a/lib/__tests__/reply-recipients.test.ts
+++ b/lib/__tests__/reply-recipients.test.ts
@@ -124,6 +124,40 @@ describe('buildReplyRecipients', () => {
     });
   });
 
+  describe('message from another connected account', () => {
+    // Two accounts connected: book@ sent this to mail@, and we reply from mail@.
+    const ALL = ['mail@hotel.example', 'book@hotel.example'];
+    const THIS_ACCOUNT = ['mail@hotel.example'];
+    const fromOtherAccount = {
+      from: [{ email: 'book@hotel.example', name: 'Booking' }],
+      to: [{ email: 'mail@hotel.example' }, { email: 'guest@other.com' }],
+      cc: [{ email: 'book@hotel.example' }],
+    };
+
+    it('replies to the other account, not back to ourselves', () => {
+      const { to, cc } = buildReplyRecipients(fromOtherAccount, 'reply', ALL, THIS_ACCOUNT);
+      expect(emails(to)).toEqual(['book@hotel.example']);
+      expect(cc).toEqual([]);
+    });
+
+    it('reply-all still drops every connected account from the other recipients', () => {
+      const { to, cc } = buildReplyRecipients(fromOtherAccount, 'replyAll', ALL, THIS_ACCOUNT);
+      expect(emails(to)).toEqual(['book@hotel.example', 'guest@other.com']);
+      expect(cc).toEqual([]);
+    });
+
+    it('still continues the thread for a message this account sent', () => {
+      const ownSent = { ...fromOtherAccount, from: [{ email: 'mail@hotel.example' }], to: [{ email: 'guest@other.com' }] };
+      const { to } = buildReplyRecipients(ownSent, 'reply', ALL, THIS_ACCOUNT);
+      expect(emails(to)).toEqual(['guest@other.com']);
+    });
+
+    it('without selfSentEmails every connected account counts as self (single-account behaviour)', () => {
+      const { to } = buildReplyRecipients(fromOtherAccount, 'reply', ALL);
+      expect(emails(to)).toEqual(['mail@hotel.example', 'guest@other.com']);
+    });
+  });
+
   it('returns nothing without a source message', () => {
     expect(buildReplyRecipients(undefined, 'replyAll', OWN)).toEqual({ to: [], cc: [] });
   });
diff --git a/lib/reply-recipients.ts b/lib/reply-recipients.ts
index fdf260825..380980e00 100644
--- a/lib/reply-recipients.ts
+++ b/lib/reply-recipients.ts
@@ -95,11 +95,19 @@ export function isSelfSent(source: ReplySource | undefined, ownEmails: string[])
  *
  * A self-sent message with no visible recipients (Bcc-only) has nothing to
  * continue to, so it falls back to the normal behaviour.
+ *
+ * `selfSentEmails` decides what counts as "self-sent" and defaults to
+ * `ownEmails`. With several connected accounts, `ownEmails` holds every
+ * account's addresses (so reply-all still drops all of them), but a message
+ * one of those accounts sent to another is a received message for the
+ * account it landed in: pass only that account's addresses here, or the
+ * reply goes back to the original To - the account itself.
  */
 export function buildReplyRecipients(
   source: ReplySource | undefined,
   mode: 'reply' | 'replyAll',
   ownEmails: string[],
+  selfSentEmails: string[] = ownEmails,
 ): ReplyRecipientsResult {
   if (!source) return { to: [], cc: [] };
 
@@ -108,7 +116,7 @@ export function buildReplyRecipients(
   const replyToList = withEmail(source.replyToAddresses);
   const externalReplyTo = replyToList.filter((r) => !isOwnAddress(r.email, ownEmails));
 
-  if (!externalReplyTo.length && isSelfSent(source, ownEmails)) {
+  if (!externalReplyTo.length && isSelfSent(source, selfSentEmails)) {
     const originalTo = withEmail(source.to);
     if (originalTo.length > 0) {
       return {